Well, there's lots of reasons. Only issue is that I'm having trouble finding/coming up with a definitive list to bring up when I get asked this question or when a 1DsIII vs 5DII comparison comes up. There's the obvious big-box features (My 1D4 shoots 2 FPS faster than your 7D!/My DsIII has 100% viewfinder coverage), but there's all the 'intangibles' that aren't listed on a Dxomark or Snapsort comparison.
Here's what I have so far:
Reduced shutter lag
Higher flash sync speed
Weathersealing/build/vertical grip
45-point AF system
Higher AF motor drive speed due to higher voltage batteries
Silent-shooting mode
Higher quality support circuity (see 5DII's banding in shadow compared to the DsIII's, despite being on the same sensor)
Increased customization available through personal/custom functions
What else?

I've already heard complaints from those who've tried my 50D with my 28mm, and it's a lightweight compared to the 1D series.



