Nothing!

The mark is really intended to accurately measure the amount of lens extension you're using when using extension tubes and bellows, so that you can look up the amount of exposure compensation needed to make allowance for light fall-off (due to the extension), or to accurately measure film plane to subject distance. There are exposure compensation tables available which tell you the amount of compensation required for a given amount of lens extension, too.
If you're more mathematically inclined you can work out the exposure compensation yourself using the Inverse Square Law ...
http://csep10.phys.utk.edu …lect/light/intensity.html
Thanks. Exposure compensation never entered my mind. But if that is a consideration, then Frank (PhotosGuy) did have a point about the histogram since one can use it to check exposure, too. But then, Frank likes using histograms to using a handheld light meter, too. 



