It focus more faster (with 400mm f5.6), but it also hunt like before if you don't have a contrasty subject. In dense bush with small bird I encounter some erratic AF in servo mode. Every time a small branche move around I lost the focus point, so the AF are more sentive but with drawback. The tracking in open area are marvelous, the much bigger improvement are when subject are coming to you, I shoot a squirell (at 50-75ft away) coming to me with 100% rate. Unfortunatly when I look at my picture in the computer my smile go down. The squirrel was on an electrical cable against a grey sky, so a contrasty scene, at 400ISO i've got some chroma noise in the shadow part of the animal, bad point! Also in less than good condition (contrasty scene, shadow, etc) the final image are whooly with artifact and even if I took off the sharpness to do it later in photoshop the final looklike a point and shoot digital shot. This only happen with the 400mm, with other lens the render are marvelous (even with the ef-s 17-85). 400L USM not on par with the 40D? Problem resolving resolution of long lens shot?