The 70-300 is an impressive lens for the money (keep that in mind):
- Highest MTF in it's class (which, you will note, is different from the 70-200...)
- Gives you a stabilization system that is top notch (Tamron is known to have a killer IS system, so long as you don't plan on panning...)
- Has the new USD AF (which is SILENT, but not as fast as one would think)
- Will cost you - after rebate - $400.00. Only!
I'd be willing to wager that the Canon will focus more quickly, but if you are shooting subjects that are mostly staying still in darker conditions, than that shouldn't bother you. If you are shooting subjects that are mostly staying still in darker conditions, you also shouldn't be too bothered by the lower max. aperture, as the great VC will cover you there as well.
My vote would be - assuming moving subjects in low light aren't your primary target - for the Tamron because of the extra versatility and lower cost.